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Stacey Harmer


Plant Biology

2129 Life Sciences
(530) 752-8101


1998 PhD University of California, San Francisco Biochemistry
1991 BA University of California, Berkeley Biochemistry

Research Interests:

Understanding the molecular basis of circadian rhythms in plants and how these daily rhythms affect plant physiology.


American Society of Photobiology - New Investigator Award, 2004
Chancellor’s Fellow, UC Davis, 2011-2016

Department and Center Affiliations:

Department of Plant Biology

Professional Societies:

American Society of Plant Biologists
Society for Research on Biological Rhythms

CBS Graduate Group Affiliations:

Plant Biology  

Specialties / Focus:

Model Plants

Plant Biology
Cell and Developmental Biology
Molecular Biology, Biochemistry, and Genomics

Graduate Groups not Housed in CBS:

Designated Emphasis in Biotechnology  


Last updated 12/14/2013
Hsu, P.Y., Devisetty, U.K., and Harmer, S.L. (2013) Cycling activators control the evening arm of the plant clock. eLife. 2:e00473. doi: 10.7554/eLife.00473.

Zemach, A., Kim, M.Y., Hsieh, P.-H., Coleman-Derr,D., Eshed-Williams, L., Thao, K, Harmer, S.L., and Zilberman, D. (2013) The Arabidopsis nucleosome remodeler DDM1 allows DNA methyltransferases to access H1-containing heterochromatin. Cell 153(1):193-205. doi: 10.1016/j.cell.2013.02.033.

Hu, W., Franklin, K.A., Sharrock, R.A., Jones, M.A., Harmer, S.L., and Lagarias, J.C. (2013) Unanticipated regulatory roles for Arabidopsis phytochromes revealed by null mutant analysis. Proceedings of the National Academy of the Sciences, 110(4) 1542-1547. doi: 10.1073/pnas.1221738110

Hsu, P.Y. and Harmer, S.L. (2012) Circadian phase has profound effects on differential expression analysis. PLoS One, 7(11):e49853. doi: 10.1371/journal.pone.0049853.

Jones, M.A., Williams, B.A., McNicol J., Simpson, C.G., Brown, J.W., and Harmer, S.L. (2012) Mutation of Arabidopsis SPLICEOSOMAL TIMEKEEPER LOCUS 1 causes circadian clock defects. Plant Cell, 24(10):4066-82. doi: 10.1105/tpc.112.104828.

Nozue, K., Harmer. S.L., and Maloof, J.N. (2011) Genomic analysis of circadian clock-, light-, and growth-correlated genes reveals PIF5 as a modulator of auxin signaling in Arabidopsis. Plant Physiology, 156(1): 357-372.

Rawat, R., Takahashi, N., Hsu, P.Y., Jones, M.A., Schwartz, J., Salemi, M.R., Phinney, B.S., and Harmer, S.L. (2011) REVEILLE8 and PSEUDO-REPONSE REGULATOR5 form a negative feedback loop within the Arabidopsis circadian clock. PLoS Genetics, 7(3): e1001350.

Kerwin, R.E., Jimenez-Gomez, J.M., Fulop, D., Harmer, S.L., Maloof, J.N., Kliebenstein, D.J. (2011) Network QTL mapping of circadian clock outputs identifies metabolic pathway to clock linkages in Arabidopsis. Plant Cell, 23(2):471-85.

Jones, M.A., and Harmer, S.L. (2011) JMJD5 functions in concert with TOC1 in the Arabidopsis circadian system. Plant Signaling & Behavior 6(3): 1-4.

Ellison ,C.T., Vandenbussche, F., Van Der Straeten, D., Harmer, S.L. (2010) XAP5 CIRCADIAN TIMEKEEPER regulates ethylene responses in aerial tissues of Arabidopsis. Plant Physiology, 2010 Dec 16. [Epub ahead of print] PubMed PMID: 21163961.

Jones, M.A., Covington, M.F., Ditacchio, L., Vollmers, C., Panda, S., Harmer, S.L. (2010) Jumonji domain protein JMJD5 functions in both the plant and human circadian systems. Proceedings of the National Academy of the Sciences 107(50): 21623-21628.

Harmer, S.L. (2010). Plant biology in the fourth dimension. Plant Physiology, 154:467 - 470.

Rawat, R., Schwartz, J., Jones, M.A., Sairanen, I., Cheng, Y., Andersson, C.R., Zhao, Y., Ljung, K., and Harmer, S.L. (2009). REVEILLE1, a Myb-like transcription factor, integrates the circadian clock and auxin pathways. Proceedings of the National Academy of the Sciences 106(39) 16883-16888.

Laboratory Personnel:

Harmer Lab

Teaching Interests:

Plant biology, molecular biology, genetics, signal transduction. http://


MCB 121 Mol Bio Eukaryotic Cells Fall
MCB 160L Genetics Lab Winter